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MONITORING NESTS
Active nests must be checked for fate every 3--4 days, and ideally every
other day. The longer the interval between checks the more difficult it is
to determine the fate of the nest. See Martin and Geupel (1993) for more
information on monitoring nests. The information from routine monitoring of
nests is used to determine nest fate, productivity, causes of failure and to
calculate variables needed for Mayfield nesting success estimates. New
participants should request examples of nest cards to help understand the
importance of good data reporting. An entire nest visit record line should
be filled in each time a nest is visited, even if no activity is observed.
Record the number of eggs or young at each visit if you count them even if
the number has not changed since the last visit. It is critical to check
nests often and to record detailed descriptions of everything you
see during each visit.
There are several guidelines that should be followed, in order to
minimize disturbance of nests while still providing comprehensive
information about a nest and it's fate.
Nest Check Guidelines
- Check nests from a distance whenever possible, and don't disturb the
birds unless a transition from one stage of nesting to the next is expected.
Don't go more than 8 days without actually checking nest contents.
- Check nests more frequently (every 2 days) when transitions (onset of
INCUBATION, HATCH, FLEDGING) are expected or when you don't have enough
data to estimate when hatch or fledging might occur. The tighter your
visits bracket these events the better the data.
- Don't disturb birds during building or early in egg-laying because
they may abandon.
- Keep careful track of the stage of your nests. Use a table of clutch
sizes, and incubation and nestling periods to estimate when incubation,
hatch, and fledging may occur so you can plan when more frequent nest
visits are needed.
- Avoid creating dead end paths when checking nests. Approach the nest
along one route and leave by another. This makes it more difficult for
predators to determine exact nest locations from watching your
activity or following scent trails.
- Never go near an active nest if predators are nearby or watching you.
Determining Nesting Stage -- Useful Cues
Even if you cannot see the contents of a nest, there are still many
useful cues that can be used to determine the stage of the nest in the
nesting cycle. Be particularly conscious of evidence of nest building or
egg-laying and do not approach nests at these times. The following are
particularly useful information:
- Adults with building material
- This almost certainly indicates that birds are still building their
nest. Be careful not to get too close to the nest during building -- the
birds may abandon.
- Mate-feeding
- males of some open- and cavity-nesting species will feed
the female on the nest during egg laying, incubation, and early brood
rearing. A parent going to a nest with food may actually be feeding its
mate, not nestlings. Be careful in interpreting feeding when you only
observed one parent. Don't jump to conclusions, simply write down what
you see. It is particularly difficult to recognize mate-feeding as opposed
to nestling feeding in cavity-nesters. Pairs of many species will exchange
contact calls during mate-feeding. Sometimes you can see an adult stick
its head out of a cavity to get the food. Note: adults will occasionally
enter the nest hole to mate feed and nestlings will also stick their heads
out for food late in the nestling period.
- Fecal Sacs
- parents will carry fecal sacs away from the nest during the
early and middle parts of the nestling period.
- Feeding Young
- in cavity-nesters, parents will go into the cavity to feed
the young until the end of the brood-rearing period. A parent feeding
from outside early in brood-rearing is often an indication that the other
parent is in the cavity. Always record whether parents are feeding from
the inside or outside of cavities and give a best guess as to whether the
one accepting food is an adult or young and why you guess that.
- Noisy Babies
- Young of many species are very noisy, especially late in the
nestling period (e.g. woodpeckers, sapsuckers, orioles). This can be
useful in finding nests, and in ageing the young. Listen carefully for
noise from nests and record distance at which it is audible. Once noise
has been heard, record lack of noise as well.
Filling Out The Nest Card
As in the previous section, UNDERLINED VARIABLES IN LARGE BOLD
LETTERS are included in the BBIRD database. The variable codes
in parentheses after each variable are the exact names that should
be used in data files contributed to the national database.
Underlined Variables in italics and mixed case are needed to
determine nest fate and/or provide useful information. Italicized
variables are not reported directly to the central data repository but
some are needed to calculate BBIRD reporting variables. In Arizona we
reserve most of the space on the backs of our nest cards for recording
information from visits to nests:
FIGURE: NEST VISIT INFORMATION (TOP BACK OF NEST CARD)
At each nest visit the date and time of day are recorded. Then the
following information is written on the field nest card:
- #Eggs
- record a value here only when the nest contents are actually viewed.
All nests within 6 m of the ground should be checked by eye or with a
mirror pole once during incubation (early is best). Two sections of half
inch PVC with a shaving mirror or automobile rear-view mirror on top
make a good nest monitoring pole for higher nests. Use binoculars to
look at the mirror to check the nest. A short section of larger diameter
PVC pipe or a pipe connector fitting can be glued to the end of one of the
PVC poles to act as a sleeve that will connect the 2 poles when in use.
Extendable light bulb replacement and paint poles can be found at some
hardware stores, and window washer poles are very good for high nests.
Nest checking poles are a pain to carry around in the field, so given an
option, field assistants will tend to forget to use them. Having multiple
poles and pushing assistants to get the data will improve your results. The
number of eggs is reported once in the BBIRD database as CLUTCH
SIZE (CS) or NON-FINAL CLUTCH SIZE (NFCS).
- #Yng
- number of hatchlings in the nest. Record a number only when actually
counted. Check once soon after hatch to get egg hatching success, and
once 2-3 days before expected fledging date to get an estimate of number
fledged. The closer your check is to fledging the better the estimate of
number fledged, but be careful of force fledging by getting too close.
Early fledging may reduce survival. Try to check from a distance,
particularly late in the nestling period. These data are used to determine
NUMBER FLEDGED (NFLDG) which is reported in the BBIRD
database.
- Elusiveness of parents
- How the adult acts around the nest before becoming
disturbed. Rate the conspicuousness or elusiveness of the parent around
the nest (within a 10 m radius). We are trying to determine if we are just
finding nests of the obvious birds, and if nest success differs between
obvious and elusive birds. Rate the elusiveness of the more conspicuous
parent that is actively involved in the primary activity of that stage of the
nest. For example, only females incubate in most passerines, so record
female elusiveness during incubation. Both male and female feed the
young in most species, so rate the elusiveness of whichever parent is more
conspicuous during the nestling period. Report elusiveness only when you
are not close to (i.e. within roughly 10 meters of) the nest, and when you
are not actively following adults. Elusiveness of parents
is a measure of how skittish parent birds are under circumstances in which
a potential nest predator (you) has not obviously found a nest, but
parental behavior could lead to the nest being detected. The following are
the codes used:
TABLE: ELUSIVENESS CODES
| Code |
Explanation |
| 1 |
Extremely shy and elusive. Would not approach
the nest with you nearby or was able to go to and from the nest without
being observed. |
| 2 |
Somewhat shy or elusive. |
| 3 |
Somewhat conspicuous |
| 4 |
Very conspicuous. Unconcerned or oblivious to your
presence. Went to the nest with you standing nearby |
- Minutes at the nest
- number of minutes you spent observing the nest. If no
activity is observed at the nest, minutes spent observing is critically
important in deciding whether the nest is still active. Parents are
sometimes away from an active nest for long periods of time. It is
sometimes necessary to spend 30 minutes or more to get a good idea of
whether or not the nest is active. Seeing no activity after 30 minutes of
observation is much stronger evidence that the nest is no longer active
than a 5 minute observation. You don't always have time for a 30 minute
observation but we need to know how long you did watch.
- Comments
- Comments are critically important, particularly when it's
impossible to check nest contents directly. These data are not reported to
the central data repository, but are essential to the determination of nest
fate and days under observation. The following is a list of useful things
that should be recorded:
- Nest active/failed -- If no activity is observed at a
nest, record how much effort you put in watching the nest (Watched 3 times
for 5 minutes each: no activity). When no activity is observed at a nest,
check it again in 2 days. Nest status is particularly difficult to
determine at cavity nests. Very little activity occurs at some cavity
nests during incubation. If no activity is seen for 6 days, sit and
observe the nest from a long distance for 30 minutes. Incubating birds will
often sit on a nest for 30 minute periods and, consequently, 5 minute nest
checks are not reliable for determining status. As a last resort, you can
try banging on the tree with a stick. Banging on the tree during incubation
will flush some cavity-nesters, but will just cause others to sit tight on
their nests.. NOTE: banging on the tree is not a substitute for
sitting and watching the nest. Bang on the tree only as a last resort after
watching the nest for at least 10 minutes. WARNING: banging on
old snags can bring down dead branches -- BE CAREFUL.
- Stage of nesting (BLDG, LAYing,
INCubation, NESTLing) -- Record the nesting stage whenever
you have some idea of the stage. If you are not 100% sure of the stage
then include a "?" after the stage. MOST IMPORTANTLY, always include
the cues you based your determination on. These notes are extremely important
for determining fate of the nest and changes in nesting stage, especially
for cavity and canopy nests. See Parent Activity section for behavior cues.
Be sure to record events such as hatching (young just hatched, egg shells
still present). If the nest is in the building stage, record an estimated
percent completion and/or the size of the nesting material being used --
birds tend to bring finer material late in the building stage.
- Age of young. -- Describe: size, extent of feather
development, and eye closure. (Example: chicks nickel sized, no feathers,
eyes closed). If you know or can estimate chick age include that, but
describe the babies anyway so we can develop aging guidelines for each
species. Record noise levels for woodpecker young (distance from nest tree
that noise is audible, and a subjective description: QUIET, NOISY, VERY
NOISY etc.).
- Parent activity -- can be used to determine whether the
nest is active and its stage in nesting. Record parent activity even if
you know nesting stage from checks of nest contents. Behaviors of
known-stage parents are useful in determining behaviors associated with
each stage of nesting. Include the sex and number of parents you observed,
and what they were doing. Always record observations of building
material, food, and fecal sacs. Don't record
"incubating" unless you are certain that a bird is sitting on eggs. If a
bird is sitting on the nest and you don't flush her off to confirm eggs, you
should record "sitting on nest" rather than "incubating". Birds will often
sit on nests that have already hatched, even late in the nestling stage.
Sometimes birds will even sit on nests for short periods during laying. Also,
for cavity-nesters, note whether parents are feeding from the inside or
outside of cavities, and if they go in, record how long they stay inside.
- When a nesting attempt terminates (fails or succeeds)
record the circumstances and any relevant observations. Check for shell
fragments, holes in the nest, torn up nests etc. If you are trying to
distinguish predation from fledging, look for fecal droppings on the edge
of the nest or on the ground under the nest, and flattening of the nest
edge caused by chicks perching there before fledging. Always look for
fledglings in the area to try to confirm fledging.
- Cowbird parasitism -- Look carefully for cowbird eggs
when you check nests. Cowbird eggs are mottled brown, about 20 mm long, and
quite rounded. Look for eggs that are different from the rest. Cowbird
chicks sometimes hatch before the host's eggs, and are often much larger
than host chicks. Cowbirds will remove host eggs from the nest. Cowbirds
will also lay eggs late in the incubation stage. Late Cowbird eggs usually
don't hatch because the host stops incubating after its eggs hatch. Look
for cowbird eggs under nestlings in frequently parasitized species. When
cowbirds are present take careful notes on the number and fate of cowbird
eggs/chicks and host eggs/chicks on each visit.
The Last Nest Visit
It is important insofar as possible to determine the fates of nests.
Without this information, we cannot obtain accurate estimates of failure
rates of nests, information that is essential for extrapolating to
population-level consequences for the species being studied.
FIGURE: NEST FATE INFORMATION (BOTTOM FRONT OF NEST CARD)
When a nest fails or fledges, immediately fill in the nest fate area of
the card (see figure, above). Include both your own best estimate of what
occurred, and detailed notes on the physical/behavioral evidence you used
to come to your conclusion. Write on the card when you are sure of nest
fate (observed some definitive cue) or if it is your best estimate. Fill in
the spaces for number of Brown-headed Cowbird (BHCO) eggs, eggs hatched, and
chicks fledged. Pay particular attention to writing notes about cause of
failure or evidence for fledging. Determining whether or not a nest
actually fledged young is difficult. In the absence of other cues, we
assume chicks fledged successfully if the median date between the last nest
check during which the nest was active and the final nest check when the nest
was empty was within 2 days of predicted fledging date. If the nest is
abandoned with eggs or chicks still in the nest, or the nest fell out of its
tree, then be sure to comment on weather during the period since the last
check (rain, cold weather, wind storms that could cause failure). Even if
the weather was good, record this fact so the person making the final
determination of nest fate can be sure you considered weather as a factor.
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