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MONITORING NESTS
Field Protocol Menu
Program Overview
Where To Look
Methods
  Data Sheets
  Establishing Study Plots
  Finding New Nests
  Monitoring Nests
  Summarizing Nest Fate
  Describing Nests' Locations
  Measuring Vegetation
  Point Counts
  Measuring Weather
References
Appendix of BBIRD Variables
General Protocol (PDF) 
Grassland Protocol (PDF) 

Active nests must be checked for fate every 3--4 days, and ideally every other day. The longer the interval between checks the more difficult it is to determine the fate of the nest. See Martin and Geupel (1993) for more information on monitoring nests. The information from routine monitoring of nests is used to determine nest fate, productivity, causes of failure and to calculate variables needed for Mayfield nesting success estimates. New participants should request examples of nest cards to help understand the importance of good data reporting. An entire nest visit record line should be filled in each time a nest is visited, even if no activity is observed. Record the number of eggs or young at each visit if you count them even if the number has not changed since the last visit. It is critical to check nests often and to record detailed descriptions of everything you see during each visit.

There are several guidelines that should be followed, in order to minimize disturbance of nests while still providing comprehensive information about a nest and it's fate.


Nest Check Guidelines

  1. Check nests from a distance whenever possible, and don't disturb the birds unless a transition from one stage of nesting to the next is expected. Don't go more than 8 days without actually checking nest contents.
  2. Check nests more frequently (every 2 days) when transitions (onset of INCUBATION, HATCH, FLEDGING) are expected or when you don't have enough data to estimate when hatch or fledging might occur. The tighter your visits bracket these events the better the data.
  3. Don't disturb birds during building or early in egg-laying because they may abandon.
  4. Keep careful track of the stage of your nests. Use a table of clutch sizes, and incubation and nestling periods to estimate when incubation, hatch, and fledging may occur so you can plan when more frequent nest visits are needed.
  5. Avoid creating dead end paths when checking nests. Approach the nest along one route and leave by another. This makes it more difficult for predators to determine exact nest locations from watching your activity or following scent trails.
  6. Never go near an active nest if predators are nearby or watching you.


Determining Nesting Stage -- Useful Cues

Even if you cannot see the contents of a nest, there are still many useful cues that can be used to determine the stage of the nest in the nesting cycle. Be particularly conscious of evidence of nest building or egg-laying and do not approach nests at these times. The following are particularly useful information:

Adults with building material
This almost certainly indicates that birds are still building their nest. Be careful not to get too close to the nest during building -- the birds may abandon.
Mate-feeding
males of some open- and cavity-nesting species will feed the female on the nest during egg laying, incubation, and early brood rearing. A parent going to a nest with food may actually be feeding its mate, not nestlings. Be careful in interpreting feeding when you only observed one parent. Don't jump to conclusions, simply write down what you see. It is particularly difficult to recognize mate-feeding as opposed to nestling feeding in cavity-nesters. Pairs of many species will exchange contact calls during mate-feeding. Sometimes you can see an adult stick its head out of a cavity to get the food. Note: adults will occasionally enter the nest hole to mate feed and nestlings will also stick their heads out for food late in the nestling period.
Fecal Sacs
parents will carry fecal sacs away from the nest during the early and middle parts of the nestling period.
Feeding Young
in cavity-nesters, parents will go into the cavity to feed the young until the end of the brood-rearing period. A parent feeding from outside early in brood-rearing is often an indication that the other parent is in the cavity. Always record whether parents are feeding from the inside or outside of cavities and give a best guess as to whether the one accepting food is an adult or young and why you guess that.
Noisy Babies
Young of many species are very noisy, especially late in the nestling period (e.g. woodpeckers, sapsuckers, orioles). This can be useful in finding nests, and in ageing the young. Listen carefully for noise from nests and record distance at which it is audible. Once noise has been heard, record lack of noise as well.


Filling Out The Nest Card

As in the previous section, UNDERLINED VARIABLES IN LARGE BOLD LETTERS are included in the BBIRD database. The variable codes in parentheses after each variable are the exact names that should be used in data files contributed to the national database. Underlined Variables in italics and mixed case are needed to determine nest fate and/or provide useful information. Italicized variables are not reported directly to the central data repository but some are needed to calculate BBIRD reporting variables. In Arizona we reserve most of the space on the backs of our nest cards for recording information from visits to nests:

FIGURE: NEST VISIT INFORMATION (TOP BACK OF NEST CARD)

At each nest visit the date and time of day are recorded. Then the following information is written on the field nest card:

#Eggs
record a value here only when the nest contents are actually viewed. All nests within 6 m of the ground should be checked by eye or with a mirror pole once during incubation (early is best). Two sections of half inch PVC with a shaving mirror or automobile rear-view mirror on top make a good nest monitoring pole for higher nests. Use binoculars to look at the mirror to check the nest. A short section of larger diameter PVC pipe or a pipe connector fitting can be glued to the end of one of the PVC poles to act as a sleeve that will connect the 2 poles when in use. Extendable light bulb replacement and paint poles can be found at some hardware stores, and window washer poles are very good for high nests. Nest checking poles are a pain to carry around in the field, so given an option, field assistants will tend to forget to use them. Having multiple poles and pushing assistants to get the data will improve your results. The number of eggs is reported once in the BBIRD database as CLUTCH SIZE (CS) or NON-FINAL CLUTCH SIZE (NFCS).
#Yng
number of hatchlings in the nest. Record a number only when actually counted. Check once soon after hatch to get egg hatching success, and once 2-3 days before expected fledging date to get an estimate of number fledged. The closer your check is to fledging the better the estimate of number fledged, but be careful of force fledging by getting too close. Early fledging may reduce survival. Try to check from a distance, particularly late in the nestling period. These data are used to determine NUMBER FLEDGED (NFLDG) which is reported in the BBIRD database.
Elusiveness of parents
How the adult acts around the nest before becoming disturbed. Rate the conspicuousness or elusiveness of the parent around the nest (within a 10 m radius). We are trying to determine if we are just finding nests of the obvious birds, and if nest success differs between obvious and elusive birds. Rate the elusiveness of the more conspicuous parent that is actively involved in the primary activity of that stage of the nest. For example, only females incubate in most passerines, so record female elusiveness during incubation. Both male and female feed the young in most species, so rate the elusiveness of whichever parent is more conspicuous during the nestling period. Report elusiveness only when you are not close to (i.e. within roughly 10 meters of) the nest, and when you are not actively following adults. Elusiveness of parents is a measure of how skittish parent birds are under circumstances in which a potential nest predator (you) has not obviously found a nest, but parental behavior could lead to the nest being detected. The following are the codes used:

TABLE: ELUSIVENESS CODES
Code Explanation
1 Extremely shy and elusive. Would not approach the nest with you nearby or was able to go to and from the nest without being observed.
2 Somewhat shy or elusive.
3 Somewhat conspicuous
4 Very conspicuous. Unconcerned or oblivious to your presence. Went to the nest with you standing nearby

Minutes at the nest
number of minutes you spent observing the nest. If no activity is observed at the nest, minutes spent observing is critically important in deciding whether the nest is still active. Parents are sometimes away from an active nest for long periods of time. It is sometimes necessary to spend 30 minutes or more to get a good idea of whether or not the nest is active. Seeing no activity after 30 minutes of observation is much stronger evidence that the nest is no longer active than a 5 minute observation. You don't always have time for a 30 minute observation but we need to know how long you did watch.
Comments
Comments are critically important, particularly when it's impossible to check nest contents directly. These data are not reported to the central data repository, but are essential to the determination of nest fate and days under observation. The following is a list of useful things that should be recorded:
  1. Nest active/failed -- If no activity is observed at a nest, record how much effort you put in watching the nest (Watched 3 times for 5 minutes each: no activity). When no activity is observed at a nest, check it again in 2 days. Nest status is particularly difficult to determine at cavity nests. Very little activity occurs at some cavity nests during incubation. If no activity is seen for 6 days, sit and observe the nest from a long distance for 30 minutes. Incubating birds will often sit on a nest for 30 minute periods and, consequently, 5 minute nest checks are not reliable for determining status. As a last resort, you can try banging on the tree with a stick. Banging on the tree during incubation will flush some cavity-nesters, but will just cause others to sit tight on their nests.. NOTE: banging on the tree is not a substitute for sitting and watching the nest. Bang on the tree only as a last resort after watching the nest for at least 10 minutes. WARNING: banging on old snags can bring down dead branches -- BE CAREFUL.
  2. Stage of nesting (BLDG, LAYing, INCubation, NESTLing) -- Record the nesting stage whenever you have some idea of the stage. If you are not 100% sure of the stage then include a "?" after the stage. MOST IMPORTANTLY, always include the cues you based your determination on. These notes are extremely important for determining fate of the nest and changes in nesting stage, especially for cavity and canopy nests. See Parent Activity section for behavior cues. Be sure to record events such as hatching (young just hatched, egg shells still present). If the nest is in the building stage, record an estimated percent completion and/or the size of the nesting material being used -- birds tend to bring finer material late in the building stage.
  3. Age of young. -- Describe: size, extent of feather development, and eye closure. (Example: chicks nickel sized, no feathers, eyes closed). If you know or can estimate chick age include that, but describe the babies anyway so we can develop aging guidelines for each species. Record noise levels for woodpecker young (distance from nest tree that noise is audible, and a subjective description: QUIET, NOISY, VERY NOISY etc.).
  4. Parent activity -- can be used to determine whether the nest is active and its stage in nesting. Record parent activity even if you know nesting stage from checks of nest contents. Behaviors of known-stage parents are useful in determining behaviors associated with each stage of nesting. Include the sex and number of parents you observed, and what they were doing. Always record observations of building material, food, and fecal sacs. Don't record "incubating" unless you are certain that a bird is sitting on eggs. If a bird is sitting on the nest and you don't flush her off to confirm eggs, you should record "sitting on nest" rather than "incubating". Birds will often sit on nests that have already hatched, even late in the nestling stage. Sometimes birds will even sit on nests for short periods during laying. Also, for cavity-nesters, note whether parents are feeding from the inside or outside of cavities, and if they go in, record how long they stay inside.
  5. When a nesting attempt terminates (fails or succeeds) record the circumstances and any relevant observations. Check for shell fragments, holes in the nest, torn up nests etc. If you are trying to distinguish predation from fledging, look for fecal droppings on the edge of the nest or on the ground under the nest, and flattening of the nest edge caused by chicks perching there before fledging. Always look for fledglings in the area to try to confirm fledging.
  6. Cowbird parasitism -- Look carefully for cowbird eggs when you check nests. Cowbird eggs are mottled brown, about 20 mm long, and quite rounded. Look for eggs that are different from the rest. Cowbird chicks sometimes hatch before the host's eggs, and are often much larger than host chicks. Cowbirds will remove host eggs from the nest. Cowbirds will also lay eggs late in the incubation stage. Late Cowbird eggs usually don't hatch because the host stops incubating after its eggs hatch. Look for cowbird eggs under nestlings in frequently parasitized species. When cowbirds are present take careful notes on the number and fate of cowbird eggs/chicks and host eggs/chicks on each visit.


The Last Nest Visit

It is important insofar as possible to determine the fates of nests. Without this information, we cannot obtain accurate estimates of failure rates of nests, information that is essential for extrapolating to population-level consequences for the species being studied.

FIGURE: NEST FATE INFORMATION (BOTTOM FRONT OF NEST CARD) Fate Variables on Nest Card
Field Protocol Menu
Program Overview
Where To Look
Methods
  Data Sheets
  Establishing Study Plots
  Finding New Nests
  Monitoring Nests
  Summarizing Nest Fate
  Describing Nests' Locations
  Measuring Vegetation
  Point Counts
  Measuring Weather
References
Appendix of BBIRD Variables
General Protocol (PDF) 
Grassland Protocol (PDF) 

When a nest fails or fledges, immediately fill in the nest fate area of the card (see figure, above). Include both your own best estimate of what occurred, and detailed notes on the physical/behavioral evidence you used to come to your conclusion. Write on the card when you are sure of nest fate (observed some definitive cue) or if it is your best estimate. Fill in the spaces for number of Brown-headed Cowbird (BHCO) eggs, eggs hatched, and chicks fledged. Pay particular attention to writing notes about cause of failure or evidence for fledging. Determining whether or not a nest actually fledged young is difficult. In the absence of other cues, we assume chicks fledged successfully if the median date between the last nest check during which the nest was active and the final nest check when the nest was empty was within 2 days of predicted fledging date. If the nest is abandoned with eggs or chicks still in the nest, or the nest fell out of its tree, then be sure to comment on weather during the period since the last check (rain, cold weather, wind storms that could cause failure). Even if the weather was good, record this fact so the person making the final determination of nest fate can be sure you considered weather as a factor.


Montana Cooperative Wildlife Research Unit

University of Montana


Home Page

About BBIRD

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Copyright © 1997, University of Montana
Last modified: Tuesday, 25-March-97 10:57:41 MST