Publications                                     Data and Models

Books

Mills, L.S. Conservation of Wildlife Populations: Demography, Genetics, and Management. Wiley / Blackwell Press, 2013 (2^nd Ed.)

Read a recent review of Conservation of Wildlife Populations in the Journal of Wildlife Management.

Mills, L.S., Tshering, and E. Cheng. Editors. 2013. Wildlife Research Techniques in Rugged Mountainous Asian Landscapes. Ugyen Wangchuck Institute for Conservation and Environment, Bhutan. ISBN: 978-99937-678-9-6.

Mills, L.S. Conservation of Wildlife Populations: Demography, Genetics, and Management. Wiley / Blackwell Press, 2007 (1^st Ed.)

Foresman, K.R., L.S. Mills, and Phurba. 2010. Procedures for Implementing Small Mammal Inventories in Bhutan. Publication of Ugyen Wangchuck Institute for Conservation and Environment, Bhutan.

Journal Articles (undergraduate co-authors indicated by **)

• Kumar, A.V., M. Zimova, T. E. Martin, and L. S. Mills. 2022. Contrasting seasonal effects of climate change influence density in a cold-adapted species. Global Change Biology DOI: 10.1111/gcb.16352
• Zimova, M., D. Moberg, L. S. Mills, A. J. Dietz, and A. Angerbjorn. 2022. Color molt phenology and camouflage mismatch in polymorphic populations of Arctic Fox. Biology Letters.
• Ferreira, M. S., T.J. Thurman, M. R. Jones, L. Farelo, A.V. Kumar, S.M.E. Mortimer, J.R. Demboski, L.S. Mills, P.C. Alves, J.Melo-Ferreira. IN REVIEW. The evolution of seasonal camouflage in white-tailed jackrabbits in response to past and future climates. Science
• Miranda, I, I. Giska, L. Farelo, J. Pimenta, M. Zimova, J. Bryk, L. Dalen, L. S. Mills, K. Zub, and J. Melo-Ferreira. 2021. Museomics dissects the genetic basis for adaptive seasonal colouration in the least weasel. Molecular Biology and Evolution. msab177. Read on Oxford Academic
• Zimova, M., S. T. Giery, S. Newey, J. J. Nowak, M. Spencer, L. S. Mills. 2020.  Lack of phenological shift leads to increased camouflage mismatch in mountain hares.  Proceedings of the Royal Society B: Biological Sciences, vol. 287.  DOI: Read on Oxford Academic
• Weckworth, J. K., B. W. Davis, M. Roelke-Parker, R. P. Wilkes, C. Packer, E. Eblate, M. Schwartz, L. S. Mills,.  2020. Identifying candidate genetic markers of CDV cross-species pathogenicity in African lions.  Pathogens 9(11). Read on Oxford Academic
• Ferreira, M. S., M. R. Jones, C. M. Callahan, L. Farelo, Z. Tolesa, F. Suchentrunk, P Boursot, L. S. Mills, P. C. Alves, J.M. Good, and J.Melo-Ferreira. 2020. The legacy of recurrent introgression during the radiation of hares. Systematic Biology. Read on Oxford Academic
• Kumar, A. V., M. Zimova, J. R. Sparks, and L. S. Mills. 2020. Snow-mediated plasticity does not prevent camouflage mismatch. Oecologia. Read on Oxford Academic
• Weckworth, J. K., B. W. Davis, E. Dubovi, N. Fountain-Jones, C. Packer, S. Cleaveland, M. E. Craft, E. Eblate, M. Schwartz, L. S. Mills, and M. Roelke-Parker. 2020. Cross-species transmission and evolutionary dynamics of canine distemper virus during a spillover in African lions of Serengeti National Park. Molecular Ecology:1–14. (Featured as News & Views perspective)
• Zimova, M., L. S. Barnard, B. M. Davis, A. V. Kumar, D. J. R. Lafferty, and L. S. Mills. 2020. Using remote cameras to measure seasonal molts. Ecosphere 11:1–9.
• Jones, M. R., L. S. Mills, J. D. Jensen, and J. M. Good. 2020. Convergent evolution of seasonal camouflage in response to reduced snow cover across the snowshoe hare range. Evolution:1–13.
• Jones, M. R., L. S. Mills, J. D. Jensen, and J. M. Good. 2020. The origin and spread of locally adaptive seasonal camouflage in snowshoe hares. American Naturalist 196:316–332.
• Ferreira, M. S., P. C. Alves, C. M. Callahan, I. Giska, L. Farelo, H. Jenny, L. S. Mills, K. Hackländer, J. M. Good, and J. Melo-Ferreira. 2020. Transcriptomic regulation of seasonal coat color change in hares. Ecology and Evolution 10:1180–1192.  (Cover Article)
• Gigliotti, L. C., N. D. Berg, R. Boonstra, S. M. Cleveland, D. R. Diefenbach, E. M. Gese, J. S. Ivan, K. Kielland, C. J. Krebs, A. V. Kumar, L. S. Mills, J. N. Pauli, H. B. Underwood, E. C. Wilson, and M. J. Sheriff. 2020. Latitudinal variation in snowshoe hare (Lepus americanus) body mass: A test of bergmann’s rule. Canadian Journal of Zoology 98:88–95.
• Zimova, M., A. P. K. Sirén, J. J. Nowak, A. M. Bryan, J. S. Ivan, T. L. Morelli, S. L. Suhrer, J. Whittington, and L. S. Mills. 2020. Local climate determines vulnerability to camouflage mismatch in snowshoe hares. Global Ecology and Biogeography 29:503–515.
• Lafferty, D. J. R., M. Zimova, L. Clontz, K. Hackländer, and L. S. Mills. 2019. Noninvasive measures of physiological stress are confounded by exposure. Scientific Reports 9:1–6.
• Bragina, E.V., R. Kays, A. Hody, C.E. Moorman, C.S. DePerno, and L. S. Mills.  2019.  White-tailed deer and coyote colonization: a response to Kilgo et al. (2019). Journal of Wildlife Management 83:1641-1643.
• Tempa, T., M. Hebblewhite, J. F. Goldberg, N. Norbu, T. R. Wangchuk, W. Xiao, and L. S. Mills. 2019. The spatial distribution and population density of tigers in mountainous terrain of Bhutan. Biological Conservation 238:108192.
• Bragina, E. V., R. Kays, A. Hody, C. E. Moorman, C. S. Deperno, and L. S. Mills. 2019. Effects on white-tailed deer following eastern coyote colonization. Journal of Wildlife Management 83:916–924.  (Most downloaded 2019 publication of The Wildlife Society; F1000 Prime Recommendation)
• Davis, B. M., A. V. Kumar, and L. S. Mills. 2019. A Camouflage Conundrum: Unexpected Differences in Winter Coat Color Between Sympatric Species. Ecosphere. 10(3): e02658
• Waples, R. S., R. Kays, R. J. Fredrickson, K. Pacifici, & L. S. Mills. 2018. Is the Red Wolf a Listable Unit Under the US Endangered Species Act ? Journal of Heredity. doi.org/10.1093/jhered/esy020.
• Jones, M.R., L.S. Mills, P. C. Alves, C. M. Callahan, J. M. Alves, D. J. R. Lafferty, F. M. Jiggins, J. D. Jensen, J. Melo-Ferreira, J.  M. Good. 2018.  Adaptive introgression underlies polymorphic seasonal camouflage in snowshoe hares.  Science 360:1355-1358.
• Mills, L. S., E. V. Bragina, A. V. Kumar, M. Zimova, D. J. R. Lafferty, J. Feltner, B. M. Davis, K. Hackländer, P. C. Alves, J. M. Good, J. Melo-Ferreira, A. Dietz, A. V. Abramov, N. Lopatina, and K. Fay. 2018. Winter color polymorphisms identify global hot spots for evolutionary rescue from climate change. Science 359:1033–1036.
• Zimova, M., K. Hackländer, J. M. Good, J. Melo-Ferreira, P. C. Alves and L. S. Mills. 2018 Function and underlying mechanisms of seasonal colour moulting in mammals and birds: what keeps them changing in a warming world?. Biological Reviews. doi:10.1111/brv.12405.
• Lafferty, D.J.R., A.V. Kumar, S. Whitcher, K. Hackländer, and L. S. Mills. 2017. Within-sample variation in snowshoe hare fecal glucocorticoid metabolite measurements. Conservation Physiology 5:1-7. 
• Ferreira, M. S., P. C. Alves, C. M. Callahan, J. P. Marques, L. S. Mills, J. M. Good, J. Melo-Ferreira. 2017. The transcriptional landscape of seasonal coat colour moult in the snowshoe hare. Molecular Ecology. 26:4173-4185.
• Cheng, E., K. E. Hodges, R. Sollmann, and L. S. Mills. 2017. Genetic sampling for estimating density of common species.  Ecology and Evolution. 7:6210–6219 [DOI: 10.1002/ece3.3137]
• Kumar, A. V., J. R. Sparks, L. S. Mills.  2017. Short-term response of snowshoe hares to western larch restoration and seasonal needle drop.  Restoration Ecology doi: 10.1111/rec.12533
• Mills, L. S.  2017.  Some matchmaking advice when translocated immigrants are a population’s last hope.  Animal Conservation (invited commentary) 20:12-13.
• Monterroso, P., G. Garrote, A. Serronha, E. Santos, M. Delibes-Mateos, J Abrantes, R Pérez de Ayala, F Silvestre, J Carvalho, I Vasco, A Lopes, E Maio, M J. Magalhães, L. S Mills,P J. Esteves, Miguel Á Simón & P C. Alves. 2016. Disease-mediated bottom-up regulation: An emergent virus affects a keystone prey, and alters the dynamics of trophic webs. Scientific Reports. DOI:10.1038/srep36072.
• Horne, J. S., J. J. Hervert, S. P. Woodroff, and L. S. Mills. 2016. Evaluating the benefit of captive breeding and reintroductions to endangered Sonoran pronghorn. Biological Conservation 196:133-146.
• Zimova, M., L. S. Mills, and J. J. Nowak. 2016. High fitness costs of climate change induced camouflage mismatch in a seasonally colour moulting mammal. Ecology Letters. doi: 10.1111/ele.1256.
• Goldberg, J. F., T. Tempa, N. Norbu, M. Hebblewhite, L. S. Mills, T. R. Wangchuk, and P. Lukacs. 2015. Examining Temporal Sample Scale and Model Choice with Spatial Capture-Recapture Models in the Common Leopard Panthera pardus. PlOS One 10:e0140757.
• Cheng, E., K.E. Hodges, and L.S. Mills.  2015.  Impacts of fire on snowshoe hares in Glacier National Park, Montna, USA.  Fire Ecology. 11(2):  119-136.
• Witczuk, J., S. Pagacz, J. Gliwicz and L.S. Mills.  2015.  Niche overlap between sympatric coyotes and bobcats in highland zones of Olympic Mountains, Washington.  Journal of Zoology.  doi: 10.1111/jzo.12270.
• Melo-Ferreira, J., F. A. Seixas, E. Cheng, L. S. Mills, and P. C. Alves. 2014. The hidden history of the snowshoe hare, Lepus americanus: extensive mitochondrial DNA introgression inferred from multilocus genetic variation. Molecular Ecology 23:4617-4630.
• Hodges, K.E., J. A. F. Cunningham, and L. S. Mills. 2014. Avoiding and escaping predators: movement tortuosity of snowshoe hares in risky habitats. Ecoscience. 21(2):  97-103.
• Cheng, E., K. E. Hodges, J. Melo-Ferreira, P. C. Alves, and L. S. Mills. 2014. Conservation implications of the evolutionary history and genetic diversity hotspots of the snowshoe hare. Molecular Ecology 23:2929-2942.
• Zimova, M., L. S. Mills, P. M. Lukacs, and M. S. Mitchell. 2014. Snowshoe hares display limited phenotypic plasticity to mismatch in seasonal camouflage. Proceedings of the Royal Society of London, Series B. 281:20140029.
• Mills, L. S., M. Zimova, J. Oyler, S. Running, J. Abatzoglou, and P. Lukacs. 2013. Camouflage mismatch in seasonal coat color due to decreased snow duration. Proceedings of the National Academy of Sciences 110:7360-7365. [widely covered by media, including National Public Radio All Things Considered (Sunday), Science News, ‘Research Highlights’ of Nature Climate Change, McClatchy Newspaper chain, Science Update, Discover Magazine; Also: “Faculty 1000” Recommendation]
• Taylor, R. L., J. D. Tack, D. E. Naugle, and L. S. Mills. 2013. Combined effects of energy development and disease on greater sage-grouse. PLOSone 8:e71256: 1-10.
• Witczuk, J., S. Pagacz, L. S. Mills. 2013. Disproportionate predation on endemic marmots by invasive coyotes. Journal of Mammalogy. 94:702-713
• Newby, J. R., L. S. Mills, T. K. Ruth, D. H. Pletscher, M. S. Mitchell, H. B. Quigly, K. M. Murphy, and R. DeSimone. 2013. Human-caused mortality influences spatial population dynamics: pumas in landscapes with varying mortality risks. Biological Conservation. 159:230-239.
• Tempa, T., M. Hebblewhite, L. S. Mills, T. Wangchuk, N. Norbu, T. Wangchuk, T. Nidup, P. Dendup, D. Wangchuk, Y. Wangdi, and T. Dorji. 2013. Royal Manas National Park, Bhutan: a hot spot for wild felids. Oryx 47:207-210.
• Hartway, C. and L. S. Mills. 2012. A meta-analysis assessing the effects of common management actions on the nest success of North American birds. Conservation Biology 26:657-666.
• Taylor, R. L., B. L. Walker, D. E. Naugle, and L. S. Mills. 2012. Managing multiple vital rates to maximize greater sage-grouse population growth. Journal of Wildlife Management 76:336-347.
• Johnson, H. E., L. S. Mills, J. D. Wehausen, T. R. Stephenson, and G. Luikart. 2011. Translating effects of inbreeding depression on component vital rates to overall population growth in endangered bighorn sheep. Conservation Biology 25:1240-1249.
• Lewis , C. W., Hodges, K. E., and L. S. Mills. 2011. Influence of stand and landscape features on snowshoe hare densities in fragmented forests. Journal of Mammalogy. 92:561-567.
• Hebblewhite, M., M. Musiani, and L. S. Mills. 2010. Restoration of gene flow among reintroduced Northern Rockies wolf populations. Molecular Ecology 19:4383-4385.
• Pierson, J. C., L. S. Mills, and D. P. Christian. 2010. Foraging patterns of cavity-nesting birds in fire-suppressed and prescribe-burned ponderosa pine forests in Montana. Open Environmental Sciences 4:49-60.
• Johnson, H. R., L. S. Mills, T. Stevenson, and J. Weyhausen. 2010. Population-specific vital rate contributions influence management of an endangered ungulate. Ecological Applications. 20:1753-1765.
• Johnson, H. R., L. S. Mills, J. Weyhausen, and T. Stevenson. 2010. Combining ground count, telemetry, and mark-resight data to infer population dynamics in an endangered species. Journal of Applied Ecology. 47:1083-1093.
• Griffin, S. C., M. L. Taper., R. Hoffman, and L. S. Mills. 2010. Ranking Mahalanobis distance models to develop specific predictions of occupancy from presence-only data. Journal of Wildlife Management. 74:1112-1121.
• **Humbert, J-Y, L. S. Mills, J. S. Horne, and B. Dennis. 2009. A better way to estimate population trends. Oikos 118:1940-1946.
• Hodges, K.. E., L. S. Mills, and K. Murphy. 2009. Distribution and abundance of snowshoe hares in Yellowstone National Park. Journal of Mammalogy. 90:870-878.
• Griffin, P. C., and L. S. Mills. 2009. Sinks without borders: snowshoe hare dynamics in a complex landscape. Oikos 118:1487-1498.
• Griffin, S.C., P. C. Griffin, M. L. Taper, and L. S. Mills. 2009. Marmots on the move? Dispersal in a declining mountain mammal. Journal of Mammalogy. 90:686-695.
• Witczuk, J., S. Pagacz, and L. S. Mills. 2008. Optimizing methods for monitoring programs: Olympic marmots as a case study. Wildlife Research 35:788-797.
• Hodges, K. E. and L. S. Mills. 2008. Designing fecal pellet surveys for snowshoe hares. Forest Ecology and Management. 256:1918-1926.
• Harris, N. C., M. Kauffman, and L. S. Mills. 2008. Inferences about ungulate population dynamics derived from age ratios. Journal of Wildlife Management 72:1143-1151.
• Griffin, S. C., M. L. Taper, R. Hoffman, and L. S. Mills. 2008. The case of the missing marmots: are metapopulation dynamics or range-wide declines responsible? Biological Conservation 141:1293-1309.
• Griffin, S. C., L. S. Mills, and M. Taper. 2007. Female Olympic marmots (Marmota Olympus) reproduce in consecutive years. American Midland Naturalist 158:221-225
• Griffin, P. C. and L. S. Mills. 2007. Pre-commercial thinning reduces snowshoe hare abundance in the short term. Journal of Wildlife Management 71:559-564.
• Griffin, S. C., T. Valois**, M. L. Taper, and L. S. Mills. 2007. Effects of tourists on behavior and demography of Olympic marmots. Conservation Biology 21:1070-1081.
• Hard, J.J., Mills, L.S. and Peek, J.M. 2006. Genetic implications of reduced survival of male red deer under harvest. Wildlife Biology 12:349-403.
• Griffin, P.C., S. C. Griffin, C. **Waroquiers, and L. S. Mills. 2005. Mortality by moonlight: predation risk and the snowshoe hare. Behavioral Ecology 16:938-944.
• Mills, L. S., P. C. Griffin, K. E. Hodges, K. McKelvey, L. Ruggiero, and T. **Ulizio. 2005. Pellet count indices compared to mark-recapture estimates for evaluating snowshoe hare density. Journal of Wildlife Management 69:1053-1062.
• Mildenstein, T. L., S. C. Stier, C. E. Nuevo-Diego, and L. S. Mills. 2005. Habitat selection of endangered and endemic large flying-foxes in Subic Bay, Philippines. Biological Conservation 126:93-102.
• Schwartz, M. K., and L. S. Mills. 2005. Gene flow after inbreeding leads to higher survival in deer mice. Biological Conservation 123:413-420.
• Tallmon, D. A., and L. S. Mills. 2004. Edge effects and isolation: California red-backed voles revisited. Conservation Biology 18:1658-1664.
• Leberg, P. L., M. Carloss, L. Dugas, K. L. Pilgrim, L. S. Mills, M. C. Green, and D. S. Scognamillo. 2004. Recent record of a cougar in Louisiana, with notes on diet, based on analysis of fecal materials. Southeastern Naturalist 2:653-658.
• Schwartz MK, Mills LS, Ortega Y, Ruggiero L, Allendorf FW. 2003. Landscape Location Affects Genetic Variation of Canada Lynx (Lynx canadensis). Molecular Ecology 12:1807-1816.
• Tallmon, D. A., E. S. Jules, N. J. **Radke, and L. S. Mills. 2003. Of mice and men and trillium: cascading effects of forest fragmentation. Ecological Applications 13:1193-1203.
• Griffin, P.C., L. Bienen, C. M. Gillin, and L. S. Mills. 2003. Estimating pregnancy rates and litter size in snowshoe hares using ultrasound. Wildlife Society Bulletin 31:1066-1072.
• **Riddle, A. E., K. L. Pilgrim, L. S. Mills, K. S. McKelvey, L. F. Ruggiero. 2003. Identification of mustelids using mitochondrial DNA and non-invasive sampling. Conservation Genetics. 4:241-243.
• Funk, W.C., and L. S. Mills. 2003. Potential causes of population declines in forest fragments in an Amazonian frog. Biological Conservation. 111:205-214.
• Hoekman, S. T., L. S. Mills, D. W. Howerter, J. H. Devries, and I. J. Ball. 2002. Sensitivity analysis of the life cycle of mid-continent mallards. Journal of Wildlife Management. 66:883-900.
• Mills, L. S. 2002. False samples are not the same as blind controls. Nature 415:471.
• McKelvey, K. S., G. W. McDaniel, L. S. Mills, P. C. **Griffin. 2002. Effects of plot size and shape on pellet density estimates for snowshoe hare. Wildlife Society Bulletin 30:751-755.
• Schwartz, M. K., L. S. Mills, K.S. McKelvey, L.F. Ruggiero, and F. W. Allendorf. 2002. DNA reveals high dispersal synchronizing the population dynamics of Canada lynx. Nature 415:520-522.
• Biek, R., W. C. Funk, B. A. Maxell, and L. S. Mills. 2002. What is missing in amphibian decline research: Insights from ecological sensitivity analysis. Conservation Biology. 16:728-734.
• Tallmon, D. A., H. M. **Draheim, L. S. Mills, and F. W. Allendorf. 2002. Insights into recently fragmented vole populations from combined genetic and demographic data. Molecular Ecology 11:699-709.
• **Biek, R., L. S. Mills, and B. Bury. 2002. Terrestrial and stream amphibians across clear-cut-forest interfaces in the Siskiyou Mountains, Oregon. Northwest Science 76:129-140.
• Reed, J. M., L. S. Mills, J. B. Dunning, Jr., E. S. Menges, K. S. McKelvey, R. Frye, S. R. Beissinger, and M-C Anstett, and P. Miller. 2002. Emerging issues in population viability analysis. Invited Paper, Conservation Biology 16:7-19.
• Mills, L. S., K. L. Pilgrim, M. K. Schwartz, and K. McKelvey. 2000. Identifying lynx and other North American felids based on mtDNA analysis. Conservation Genetics 1:285-288.
• Mills, L. S., D. F. Doak, and M. J. Wisdom. 2001. Elasticity analysis for conservation decision-making: Reply to Ehrlen et al., Conservation Biology 15:281-283.
• Mills, L. S., J. J. Citta, K. Lair, M. Schwartz, D. Tallmon. 2000. Estimating animal abundance using non-invasive DNA sampling: Promise and Pitfalls. Ecological Applications 10:283-294.
• Wisdom, M. J., L. S. Mills, and D. F. Doak. 2000. Life-stage simulation analysis: estimating vital rate effects on population growth for conservation. Ecology 81:628-641.
• Mills, L. S., D. F. Doak, and M. J. Wisdom. 1999. The reliability of conservation actions based on sensitivity analysis of matrix models. Conservation Biology 13:815-829.
• Jules, E., E. Frost, D. Tallmon, and L. S. Mills. 1999. Ecological consequences of forest fragmentation in the Klamath region. Natural Areas Journal 19:368-378.
• Citta, J. J., and L. S. Mills. 1999. What do demographic sensitivity analyses tell us about controlling brown-headed cowbirds? Studies in Avian Biology 18:121-134.
• Soulé, M. E., and L. S. Mills. 1998. No need to Isolate Genetics. Science 282:1658-1659.
• Wisdom, M. J., and L. S. Mills. 1997. Using sensitivity analysis to guide population recovery: Prairie chickens as an example. Journal of Wildlife Management 61:302-312.
• Mills, L. S., and F. W. Allendorf. 1996. The one-migrant-per-generation rule in conservation and management. Conservation Biology 10:1509-1518.
• Mills, L. S. 1996. “Cheetah extinction: genetics or extrinsic factors?” Conservation Biology 10:315.
• Morrison, M. L., L. S. Mills, and A. J. Kuenzi. 1996. Study and management of an isolated, rare population: The Fresno Kangaroo rat. Wildlife Society Bulletin 24:602-606.
• Power, M.E., D. Tilman, J. Estes, B. A. Menge, W. J. Bond, L. S. Mills, G. Daily, J. C. Castilla, J. Lubchenco, and R. T. Paine. 1996. Challenges in the quest for keystones. Bioscience 46:609-620.
• Mills, L. S., S. G. Hayes, C. Baldwin, M. J. Wisdom, J. Citta**, D. J. Mattson, and K. Murphy. 1996. Factors leading to different viability predictions for a grizzly bear data set. Conservation Biology 10:863-873.
• Mills, L. S. 1995. Edge effects and isolation: red-backed voles on forest remnants. Conservation Biology 9:395-403.
• Power, M. E., and L. S. Mills. 1995. The keystone cops meet in Hilo. Trends in Research in Evolution and Ecology. 10:182-184.
• Scott, J. M., T. H. Tear, and L. S. Mills. 1995. Socioeconomics and the recovery of endangered species: biological assessment in a political world. Conservation Biology 9:214-216.
• Mills, L. S., and P. E. Smouse. 1994. Demographic consequences of inbreeding in remnant populations. American Naturalist 144:412-431.
• Doak, D., and L. S. Mills. 1994. A useful role for theory in conservation. Ecology 75:615-626.
• **Clarkson, D. A., and L. S. Mills. 1994. Ecological factors associated with Hypogeous sporocarps in fragmented forests. Northwest Science 68:259-265.
• **Tallmon, D.A., and L. S. Mills. 1994. Log use and home ranges of California red-backed voles on a forest remnant. Journal of Mammalogy. 75:97-101.
• Mills, L. S., M. S. Soulé, and D. F. Doak. 1993. The keystone species concept in ecology and conservation. Bioscience 43:219-224.
• Mills, L. S., R. J. Fredrickson, B. B. Moorhead. 1993. Characteristics of old-growth forests associated with northern spotted owls in Olympic National Park. Journal of Wildlife Management 57:315-321.
• Mills, L. S., and F. F. Knowlton. 1991. Coyote space use in relation to prey abundance. Canadian Journal of Zoology 69:1516-1521.
• Mills, L. S., and F. F. Knowlton. 1989. Observer performance in known and blind radio-telemetry accuracy tests. Journal of Wildlife Management. 52:340-342.

Articles in Books (and Book Reviews in Journals) (All invited articles):

• Horne, J. S., L. S. Mills, K. M. Strickler, S. A. Temple, and J.M. Scott. 2018. Ecology and management of small populations.  Pages XX-YY In Techniques for wildlife investigations and management (ed. N. Silvy).  Seventh edition.  The Wildlife Society, Bethesda, Maryland, USA.
• Mills, L. S. 2016.  "Adaptive responses in animals to climate change" AccessScience/McGraw-Hill Education. DOI: http://accessscience.com/content/YB160512
• Mills, L. S., and H. E. Johnson. 2013. Wildlife Population Dynamics. In Krausman, P., editor. Wildlife management: Contemporary principles and practices. John Hopkins University Press.
• Mills, L. S., J. M. Scott, K. M. Strickler, and S. A. Temple. 2012. Ecology and management of small populations. Pages 270-292 In Techniques for wildlife investigations and management (ed. N. Silvy). Sixth edition. The Wildlife Society, Bethesda, Maryland, USA.
• Kelly, M. J., J. Betsch, C. Wultsch, B. Mesa, and L. S. Mills. 2011. Noninvasive Sampling for carnivores. Pages 47- 69 In Boitani, L. and R. Powell, editors. Carnivore Ecology and Conservation. Oxford University Press, Oxford, UK.
• Running, S. W. and L. S. Mills. 2009. Terrestrial Ecosystem Adaptation. Resources for the Future Report for the Climate Policy Program at RFF. Access at: http://www.rff.org/rff/documents/RFF-Rpt-Adaptation-RunningMills.pdf
• Mills, L. S. 2008. Crossing Disciplines for Endangered Species. Review for Ecology of Scott, J. M., D. D. Goble, and F.W. Davis, editors. The Endangered Species Act at Thirty: Conserving Biodiversity in Human-Dominated Landscapes (Volume 2). Island Press, Washington, D.C. Ecology 89:592-593.
• Mills, L. S. 2006 “Contributing Author” to “Chapter 14: North America” by Field, C. B., L. D. Mortsch, M. Brklacich, D. L. Forbes, P. Kovaks, J. A. Patz, S. W. Running and M. J. Scott, in: Climate Change 2007: Impacts, Adaptation and Vulnerability. Contribution of Working Group II to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change, M. L. Parry, O. F. Canziani, J. P. Palutikof, P. J. van der Linden and C. E. Hanson, Eds., Cambridge University Press, Cambridge, UK, 6127-652.
• Mildenstein, T.L., L.S. Mills, P. Landres, G. Rochefort, J. Scharberl. 2006. Recreation impacts on wildlife: review of recent research and a call for investigation of population and community level effects. Final report to Mount Rainier National Park.
• Mills, L. S., J. M. Scott, K. M. Strickler, and S. A. Temple. 2005. Ecology and management of small populations. Pages 691-713 in Braun, C. E., editor. Techniques for wildlife investigations and management. Sixth edition. The Wildlife Society, Bethesda, Maryland, USA. (Invited Article)
• Mills, L. S., and M. E. Soulé. 2006. A brief history of the role of genetics in conservation. Invited article (“box”) in Conservation Genetics textbook by F. Allendorf and G. Luikart.
• Mills, S., M. K. Schwartz, D. A. Tallmon, and K. P. Lair. 2003. Measuring and interpreting changes in connectivity for mammals in coniferous forests. Pages 587-613 in C. J. Zabel and R. G. Anthony, editors. Mammal Community Dynamics: Management and Conservation in the Coniferous Forests of Western North America. Cambridge University Press, New York, USA.
• Griffin, P. C., and L. S. Mills. 2003 “Snowshoe hares in a dynamic managed landscape.” Pages 438-449 in Editors H. R. Akcakaya, M. A. Burgman, O. Kindvall, C. Wood, P. Sjogren-Gulve, J. Hatfield, and M. A. McCarthy. Species Conservation and Management: Case Studies. Oxford University Press.
• Mills, L. S., and M. Lindberg. 2002. “Sensitivity Analysis to Evaluate the Consequences of Conservation Actions.” Pages 338-366 in R. Beissinger and D. R. McCullough, editors. Population Viability Analysis. University of Chicago Press.
• Mills, L. S. 2002. “Genetics, Demography, and Viability of Fragmented Populations.” (Book Review) Quarterly Review of Biology 77:222-223.
• Mills, L. S., and D. Tallmon. 1999. “Genetic issues in forest fragmentation.” Pages 171-184 in Forest Fragmentation: Wildlife and Management Implications. Rochelle, L. A. Lehmann, and J. Wisniewski, eds. Brill Publishers (Netherlands).
• Dobson, A., K. Ralls, M. Foster, M. E. Soule, D. Simberloff, D. Doak, J. A. Estes, L. S. Mills, D. Mattson, R. Dirzo, H. Arita, S. Ryan, E. A. Norse, R. F. Noss, and D. Johns. 1999. “Connectivity: maintaining flows in fragmented landscapes.” Pages 129-171 In Soulé, M. E. and J. Terborgh, editors. Continental Conservation: Scientific Foundations of Regional Reserve Networks. Island Press.
• Groom, M., D. B. Jensen, R. L. Knight, S. Gatewood, L. Mills, D. Boyd-Heger, L. S. Mills, and M. E. Soulé. 1999. “Buffer zones: benefits and dangers of compatible stewardship.” Pages 191-198 In Soulé, M. E. and J. Terborgh, editors. Continental Conservation: Scientific Foundations of Regional Reserve Networks. Island Press.
• Mills, L. S. 1997. “Book Review: Population Management for Survival and Recovery: Analytical Methods and Strategies in Small Population Conservation.” Journal of Wildlife Management 61:251-252.
• Mills, L. S. 1996. “Fragmentation of a natural area: Dynamics of isolation for small mammals on forest remnants.” Pages 199-219 In Wright, G., editor. National Parks and Protected Areas: Their Role in Environmental Protection. Blackwell Press.
• Mills, L.S. 1996. “Keystone Species.” in Paehlke, R., editor. Encyclopedia of Conservation and Environmentalism. Garland Publishing Co., New York.
• Mills, L.S. 1995. “Keystone Species.” Pages 381-387 in Encyclopedia of Environmental Biology. Academic Press, Inc.
• Mills, L.S. 1994. “Book Review: Principles of Conservation Biology.” Northwest Science 68:303-304.
• Soulé, M.E., and L. S. Mills. 1992. “Conservation genetics and conservation biology: a troubled marriage.” Pages 55-69 in Sandlund, O. T., K. Hindar, and A. H. D. Brown, eds., Conservation of Biodiversity for Sustainable Development. Scandinavian Univ. Press, Oslo.

Other Professional Publications:

• Mills, L. S. (contributing co-author) to “Biodiversity in High Mountain Areas”, Chapter 2:  Intergovernmental Panel on Climate Change (IPCC) Special Report on the Ocean and Cryosphere in a Changing Climate. In press (2019.
• Mills, L. S.   2016. A deadly game of hide and seek.  “Crown of the Continent and the Greater Yellowstone” magazine, Spring 2016.
• Mills, L. S. 2017. Reprinting of “A deadly game of hide and seek”.  In Mountain Views Chronicles of the Consortium for Integrated Climate Research in Western Mountains CIRMOUNT Volume 11, No. 1, May 2017 www.fs.fed.us/psw/cirmount/.
• McKelvey, K.S.; Perry, R.W.; Mills, L.S. 2013. The Effects of Climate Change on Mammals. U.S. Department of Agriculture, Forest Service, Climate Change Resource Center. http://www.fs.fed.us/ccrc/topics/wildlife/mammals/index.shtml
• Hodges, K. E. and L. S. Mills. 2005. Snowshoe hares in Yellowstone. Yellowstone Science 13:3-6.
• Yale, R. and L. S. Mills. 2000. “Do highways fragment small mammal populations?” Proceedings of the International Conference on Ecology and Transportation.
• Zager, P., L. S. Mills, W. Wakkinen, and D. Tallmon. 1995. “Woodland caribou –a conservation dilemma.” Endangered Species Update 12:1-4.
• Mills, L. S. 1995. “The use of population ecology approaches to assess re-introduction of endangered Selkirk Caribou.” Final report, submitted to: a) Idaho Fish and Game; b) Washington Department of Wildlife.
• Mills, L. S. 1995. “The role of population ecology in guiding wildlife translocations.” Final report, submitted to U.S. Forest Service, Missoula, MT.
• Cassidy, K., E. O. Garton, W. B. Krohn, L. S. Mills, J. M. Scott, and K. Williams. 1994. “National guidelines for assessment of reliability of GAP vertebrate distributions.” National Biological Survey.
• Mills, L. S. 1994. “Effects of forest fragmentation on small mammals in Southwest Oregon.” COPE Report [(Coastal Oregon Productivity Enhancement Program), an educational non-technical publication for forest managers.] 7:6-8.
• Mills, L. S. and M. Morrison. 1993. “Final Report: Ecology of the Fresno Kangaroo Rat and associated small mammals at NAS Lemoore”. Prepared for Lemoore Naval Air Station.
• Fredrickson, R.J., L.S. Mills, and B.B. Moorhead. 1989. “Spotted owl surveys in Olympic National Park — 1988 and 1989.” Olympic National Park, Port Angeles, WA.
• Mills, L.S., R.J. Fredrickson, B.B. Moorhead, and D.U. Sharp. 1988. “Spotted owl distribution along elevation and vegetation gradients in Olympic National Park.” 1988 Progress Report. Olympic National Park, Port Angeles, WA.