2022

Martin, H., M. Hebblewhite, A. Hubbs, R. Corrigan, and E. H. Merrill. 2022. Male Elk Survival, Vulnerability and Antler Size in a Transboundary Partially Migratory Elk Population. Journal of Wildlife Management, In Review.

Steenweg, R., M. Hebblewhite, J. Whittington, A. C. Burton, N. Heim, J. T. Fisher, A. Ladle, W. Lowe, T. Muhly, J. Paczkowski, and M. Musiani. 2022. Testing umbrella species and food-web properties of large carnivores in the Rocky Mountains. Biological Conservation, In Review.

Tucker, M. A., …, M. Hebblewhite, E. H. Merrill, C. Lamb, W. Peters, …, and T. Mueller. 2022. Changes in human presence during COVID-19 lockdowns altered the spatial behavior of terrestrial mammals. Science, In Revision.

Williams, S., M. Hebblewhite, H. Martin, J. Killeen, J. Berg, K. M. MacAulay, P. Smolko, and E. H. Merrill. 2021. Predation Risk Drives Long-Term Shifts in Migratory Behavior and Demography in Large Herbivore Population. Journal of Animal Ecology, In Revision.

Martin, H., M. Hebblewhite, and E. H. Merrill. 2022. Large herbivores in a partially migratory population search for the ideal free home. Ecology:14. https://doi.org/10.1002/ecy.3652

Whittington, J., R. Baron, M. Hebblewhite, A. T. Ford, and J. Paczkowski. 2022. Towns and trails drive carnivore connectivity using a step selection approach. Movement Ecology, 10: 17.

MacAulay, K. M., E. Spilker, J. Berg, M. Hebblewhite, and E. H. Merrill. 2022. Predicting multi-predator risk to elk (Cervus canadensis) using scats: Are migrant elk exposed to different predation risk? Ecology & Evolution, 12: 2.

Normandeau, J., C. Cassady-St.Clair, S. Kutz, M. Hebblewhite, and E. H. Merrill. 2022. What makes elk tick: winter tick grooming behavior in wild elk (Cervus canadensis). Journal of Mammalogy, 103: 373 - 382.

2021

Brandell, E. E., P. C. Cross, M. E. Craft, D. W. Smith, E. J. Dubovi, M. L. J. Gilbertson, T. Wheeldon, J. A. Stephenson, S. Barber-Meyer, B. L. Borg, M. Sorum, D. R. Stahler, A. Kelly, M. Anderson, H. D. Cluff, D. R. MacNulty, D. E. Watts, G. H. Roffler, H. Schwantje, M. Hebblewhite, K. Beckmen, and P. J. Hudson. 2021. Patterns and processes of pathogen exposure in gray wolves across North America. Scientific Reports 11:3722.

Berg JE, Reimer J, Smolko P, Bohm H, Hebblewhite M, Merrill EH (2021) Mothers’ movements: shifts in calving area selection by partially migratory elk. Journal of Wildlife Management [In Press].

Normandeau J, St.Clair CC, Kutz S, Hebblewhite M, Merrill EH (2021) What makes elk tick: winter tick grooming behavior in wild elk (Cervus canadensis). Journal of Mammalogy [In Press].

Normandeau J, Martin H, Merrill EH, Hebblewhite M (2021) Potential case of pseudo-hermaphroditism in Elk (Cervus canadensis) in Alberta, Canada. Canadian Field Naturalist 134: 241-247.

Sabal MC, Boyce MS, Charpentier CL, Furey NB, Luhring TM, Martin HW, Melnychuk MC, Srygley RB, Wagner CM, Wirsing AJ, Ydenberg RC, Palkovacs EP. 2021. Predation landscapes influence migratory prey ecology and evolution. Trends Ecol Evol. 13:00124-5.

Suraci JP, Gaynor KM, Allen ML, Alexander P, Brashares JS, Cendejas-Zarelli S, Crooks K, Elbroch LM, Forrester T, Green AM, Haight J, Harris NC, Hebblewhite M, Isbell F, Johnston B, Kays R, Lendrum PE, Lewis JS, McInturff A, McShea W, Murphy TW, Palmer MS, Parsons A, Parsons MA, Pendergast ME, Pekins C, Prugh LR, Sager-Fradkin KA, Schuttler S, Şekercioğlu ÇH, Shepherd B, Whipple L, Whittington J, Wittemyer G, Wilmers CC (2021) Disturbance type and species life history predict mammal responses to humans. Global Change Biology, Early View.

Kauffman MJ, Cagnacci F, Chamaillé-Jammes S, Hebblewhite M, Hopcraft JGC, Merkle JA, Mueller T, Mysterud A, Peters W, Roettger C, Steingisser A, Meacham JE, Abera K, Adamczewski J, Aikens EO, Bartlam-Brooks H, Bennitt E, Berger J, Boyd C, Côté SD, Debeffe L, Dekrout AS, Dejid N, Donadio E, Dziba L, Fagan WF, Fischer C, Focardi S, Fryxell JM, Fynn RWS, Geremia C, González BA, Gunn A, Gurarie E, Heurich M, Hilty J, Hurley M, Johnson A, Joly K, Kaczensky P, Kendall CJ, Kochkarev P, Kolpaschikov L, Kowalczyk R, van Langevelde F, Li BV, Lobora AL, Loison A, Madiri TH, Mallon D, Marchand P, Medellin RA, Meisingset E, Merrill E, Middleton AD, Monteith KL, Morjan M, Morrison TA, Mumme S, Naidoo R, Novaro A, Ogutu JO, Olson KA, Oteng-Yeboah A, Ovejero RJA, Owen-Smith N, Paasivaara A, Packer C, Panchenko D, Pedrotti L, Plumptre AJ, Rolandsen CM, Said S, Salemgareyev A, Savchenko A, Savchenko P, Sawyer H, Selebatso M, Skroch M, Solberg E, Stabach JA, Strand O, Suitor MJ, Tachiki Y, Trainor A, Tshipa A, Virani MZ, Vynne C, Ward S, Wittemyer G, Xu W, Zuther S (2021) Mapping out a future for ungulate migrations: Limited mapping of migrations hampers conservation. Science 372:566-569.

Martin H, Hebblewhite M, Merrill EH (2021) Large herbivores in a partially migratory population search for the ideal free home. Ecology [In Revision].

Williams S, Hebblewhite M, Martin H, Killeen J, Berg J, MacAulay KM, Smolko P, Merrill EH (2021) Predation Risk Drives Long-Term Shifts in Migratory Behavior and Demography in a Large Herbivore Population. Journal of Animal Ecology [In Review].

Martin HW (2021) Ungulates Search for the Ideal Home: How Predation and Forage Differentially Affect Males and Females in a Partially Migratory Population. PhD Dissertation, University of Montana.

2020

Merrill EH, Killeen J, Pettit J, Trottier M, Martin H, Berg J, Bohm H, Eggeman S, Hebblewhite M (2020) Density-Dependent Foraging Behaviors on Sympatric Winter Ranges in a Partially Migratory Elk Population, Frontiers in Ecology and Evolution 8:269.

Normandeau J, Kutz SJ, Hebblewhite M, Merrill EH. Living with liver flukes: Does migration matter? International Journal for Parasitology: Parasites and Wildlife. 2020 May 12;12:76-84.

Aikens EO, Mysterud A, Merkle JA, Cagnacci F, Rivrud IM, Hebblewhite M, Hurley MA, Peters W, Bergen S, De Groeve J, Dwinnell SPH, Gehr B, Heurich M, Hewison AJM, Jarnemo A, Kjellander P, Kröschel M, Licoppe A, Linnell JDC, Merrill EH, Middleton AD, Morellet N, Neufeld L, Ortega AC, Parker KL, Pedrotti L, Proffitt KM, Saïd S, Sawyer H, Scurlock BM, Signer J, Stent P, Šustr P, Szkorupa T, Monteith KL, Kauffman MJ (2020) Wave-like Patterns of Plant Phenology Determine Ungulate Movement Tactics. Current Biology 30(17):3444-3449.

Hebblewhite M, Whittington J (2020) Wolves without borders: Transboundary survival of wolves in Banff National Park over three decades. Global Ecology and Conservation 24:e01293

Mahoney PJ, Joly K, Borg BL, Sorum MS, Rinaldi TA, Saalfeld D, Golden H, Latham ADM, Kelly AP, Mangipane B, Koizumi CL, Neufeld L, Hebblewhite M, Boelman NT, Prugh LR (2020) Denning phenology and reproductive success of wolves in response to climate signals. Environ. Res. Lett. 15:125001.

Weeks TC (2020) Bottom-up Herbivore-Plant Feedbacks Trump Trophic Cascades in a Wolf-Elk-Grassland System. Undergraduate Thesis. University of Montana, Missoula, MT, USA.

2019

Berg, J.E., 2019. Shifts in strategy: Calving and calf survival in a partially migratory elk population. Dissertation, Department of Biological Sciences, University of Alberta, Edmonton, Canada. 288 pages.ical Sciences, University of Alberta.
Ecology of Calf Survival Summary Report 2013 – 2016

Hessami, M. 2019. Estimating Juvenile Recruitment of Elk in an Occupancy Modeling Framework. Undegraduate Thesis. Wildlife Biology Program. University of Montana, Missoula, MT.

MacAulay, K.M. (2019) Spatial predation risk for elk (Cervus elaphus) in a multi-predator community on the Rocky Mountain East Slopes, Alberta. University of Alberta.

Berg, J.E., Hebblewhite, M., Cassady St. Clair, C. & Merrill, E.H. 2019. Prevalence and mechanisms of partial migration in ungulates. Frontiers in Ecology and Evolution, Front. Ecol. Evol. 7:325., doi: 10.3389/fevo.2019.00325

Boelman NT, Liston GE, Gurarie E, Meddens A, Mahoney PJ, Kirchner P, Bohrer G, Brinkman TJ, Cosgrove C, Eitel J, Hebblewhite M, Kimball J, Lapoint SD, Nolin A, Hojlund PS, Prugh LR, Reinking A, Vierling L (2019) Integrating snow science and wildlife ecology in Arctic-Boreal North America. Environmental Research Letters 14:010401.

Steenweg R, Hebblewhite M, McKelvey K, Lukacs P, Whittington J (2019) Species-specific trade-offs in statistical power when monitoring trends in multispecies occupancy. Ecosphere 10:e02639.

Keery L (2019) Effects of bison reintroduction on vegetation and landcover in Banff National Park. Royal Rhodes University, Advisor: Mark Hebblewhite.

2018

Metz, M. C., M. Hebblewhite, D. Emlen, D. Stahler, D. Macnulty, and D. Smith. 2018. Predation by a coursing predator shapes the evolutionary traits of ungulate weapons. Nature Ecology & Evolution 2:1619-1625.

Normandeau, J., Macaulay, K., Berg, J., Merrill, E.H. 2018. Identifying guard hairs of Rocky Mountain carnivores. Wildlife Society Bulletin, Published 21 September, 2018.

Steenweg, R., M. Hebblewhite, J. Whittington, K. Mckelvey, and P. Lukacs. 2018. Sampling scales define occupancy and the occupancy-abundance relationship in animals. Ecology 99:172-183.

Hebblewhite M, Eacker DR, Eggeman S, Bohm H, Merrill EH (2018) Density-Independent Predation Affects Migrants and Residents Equally in a Declining Partially Migratory Elk Population. Oikos 127:1304-1318.

2017

Spitz, D., Hebblewhite, M., Stephenson, T. R. "MigrateR: extending model-driven methods for classifying and quantifying animal movement behavior," Ecography, 40(6): 788-799.

Steenweg, R., Hebblewhite, M., Kays, R., Ahumanda, J., Fisher, J. T., Burton, C., Townsend, S. E., Carbone, C., Rowcliffe, J. M., Whittington, J., Brodie, J., Royle, J. A., Switalski, A., Clevenger, A.P., Heim, N., Rich, L. N. "Scaling up camera traps? Monitoring the planet's biodiversity with networks of remote sensors," Frontiers in Ecology and the Environment. 15(1): 26-34.

Tucker, M.A., Bohning-Gaese, K., Fagan, W.F., Fryxell, J.M., Van Moorter, B., Alberts, S.C., Ali, A.H., Allen, A.M., Attias, N., Avgar, T., Bartlam-Brooks, H.L.A., Bayarbaatar, B., Belant, J.L., Bertassoni, A., Beyer, D.E., Bidner, L., van Beest, F.M., Blake, S., Blaum, N., Bracis, C., Brown, D., de Bruyn, P.N., Cagnacci, F., Calabrese, J.M., Camilo-Alves, C., Chamaille-Jammes, S., Chiaradia, A., Davidson, S.C., Dennis, T., DeStefano, S., Deifenbach, D., Douglas-Hamilton, I., Fennesey, J., Fichtel, C., Fiedler, W., Fischer, C., Fischoff, I., Fleming, C.H., Ford, A.T., Fritz, S., Gehr, B., Goheen, J.R., Gurarie, E., Hebblewhite, M., Heurich, M., Hewison, A.J., Hof, C., Hurme, E., Isbell, L.A., Janssen, R., Jeltsch, F., Kaczensky, P., Kane, A., Kappler, P., Kauffman, M., Kays, R., Kimuyu, D., Koch, F., Kranstauber, B., Lapoint, S.D., Leimgruber, P., Linnell, J.D.C., Lopez-Lopez, P., Markham, A.C., Mattison, J., Medici, E.P., Mellone, U., Merrill, E.H., de Miranda-Mourao, G., Morato, R.G., Morellet, N., Morrison, T.A., Diaz-Munoz, S.L., Mysterud, A., Nandintsetseg, D., Nathan, R., Niamir, A., Odden, J., O'Hara, R.B., Olvieria-Santos, G.R., Olson, K.A., Patterson, B.D., de Paula, R.C., Pedrotti, L., Rimmler, M., Rogers, T.L., Rolandsen, C.M., Rosenberry, C.S., Rubenstein, D.I., Safi, K., Said, S., Sapir, N., Sawyer, H., Schmidt, N.M., Selva, N., Sergiel, A., Shiilegdamba, E., Silva, J.P., Singh, N.J., Solberg, E.J., et al. 2017. Moving in the anthropocene: global reductions in terrestrial mammalian movements. Science, 359, 466-469. 

Whittington, J., Hebblewhite, M., Chandler, R. 2017. Generalized spatial mark-resight models for efficient density estimation of partially marked populations: an application to grizzly bears in the Rocky Mountains. Journal of Applied Ecology, early view. doi: 10.1111/1365-2664.12954 

2016

Ahrestani, F. S., Hebblewhite, M., Smith, B., Running, S. W., Post, E.. "Variation in stability of elk and red deer populations with abiotic and biotic factors at the species-distribution scale." Ecology, 97(11): 3184-3194.

Eggeman, S., Hebblewhite, M., Bohm, H., Merrill, E.H. "Behavioral Flexibility in Migratory Behavior in a Long-lived Ungulate." Journal of Animal Ecology, 85: 785–797.

Spitz, D., Hebblewhite M., Stephenson,Thomas. "'MigrateR': Extending model-driven methods for classifying and quantifying animal movement behavior." Ecography. Nov. 2016. doi: 10.1111/ecog.02587

Steenweg, R., Hebblewhite, M., Lukacs, P., Whittington, J., Forshner, A., Johnston, B., Shepherd, B., Peterson, D. "Camera-based occupancy monitoring at large scales: Power to detect trends in grizzly bears across the Canadian Rockies," Biological Conservation, 201: 192-200. 

2013

Ahrestani, F. S., Hebblewhite, M. & Post, E. S. "The importance of observation versus process error in analyses of global ungulate populations." Scientific Reports 3, 03125, doi: 3, doi:10.1038/srep03125.

Robinson, B. G and Merrill, E. H.  "Foraging-vigilance trade-offs in a partially migratory population: Comparing migrants and residents on a sympatric range." Animal Behaviour 85: 849-856.

2012

Robinson, B.G. and E.H. Merrill. 2012. The influence of snow on the functional response of grazing ungulates. Oikos 164: 265-275.

Nelson, B., M. Hebblewhite, V. Ezenwa, T. Shury, E.H. Merrill, P.C. Paquet, F. Schmiegelow, D. Seip, G. Skinner, N. Webb. 2012. Prevalence of antibodies to canine parvovirus and distemper virus in wolves in the Canadian Rocky Mountains.  Journal Wildlife Diseases 48:68-76.

2011

Griffin, K. et al. 2011. "Neonatal mortality of elk driven by climate, predator phenology and predator diversity." Journal of Animal Ecology 80(6): 1246–1257. doi: 10.1111/j.1365-2656.2011.01856.x.

Hebblewhite, M. & Merrill, E. H. 2011. "Fitness balancing in a partially migratory ungulate through risk-forage trade-offs." Oikos, 120(12): 1860-1870.

2010

McInenly, L.E., Merrill E. H., Cahill J. F., & Juma, N. G. 2010. "Festuca campestris alters root morphology and growth at Ya Ha Tinda in response to simulated grazing and nitrogen form." Functional Ecology 24: 283-292.

Robinson, B. G., M. Hebblewhite, and E. Merrill. 2010. "Are migrant and resident elk (Cervus elaphus) exposed to similar forage and predation risk on their sympatric winter range?" Oecologia 164: 265-275.

2009 and earlier

Hebblewhite, M., & Merrill, E.H. 2009. "Trade-offs between wolf predation risk and forage at multiple spatial scales in a partially migratory ungulate." Ecology, 90(12): 3445-3454.

Hebblewhite, M., Munro, R. & Merrill, E. H. 2009. "Trophic consequences of post-fire logging in a wolf-elk system." Forest Ecology and Management, 257: 1053-1062.

Hebblewhite, M., Merrill, E. H. & McDermid, G. 2008. "A mutli-scale test of the forage maturation hypothesis for a partially migratory montane elk population." Ecological Monographs, 78: 141-166.

Hebblewhite, M., & Merrill, E.H. 2008. "Modelling wildlife-human relationships for social species with mixed-effects resource selection models." Journal of Applied Ecology, 45: 834-844.

Hebblewhite, M. 2007." Predator-prey management in the National park context: lessons from a transboundary wolf-elk, moose, and caribou system." Transactions of the North American Wildlife Conference, 72: 348-365.

Hebblewhite, M. & Merrill, E. H. 2007. "Multiscale wolf predation risk for elk: does migration reduce risk?" Oecologia, 152: 377-387.

Hebblewhite, M., Merrill, E. H., Morgantini, L. E., White, C. A., Allen, J. R., Bruns, E., Thurston, L. & Hurd, T. E. 2006. "Is the migratory behavior of montane elk herds in peril? The case of Alberta’s Ya Ha Tinda elk herd." Wildlife Society Bulletin, 34: 1280-1295

Hebblewhite, M., Merrill, E. H. & McDonald, T. E. 2005 "Spatial decomposition of predation risk using resource selection functions: an example in a wolf-elk system." Oikos 111: 101-111.

Webb, N.F., M. Hebblewhite, and E.H. Merrill. 2008. Statistical methods for identifying wolf kill sites using global positioning system locations. Journal Wildlife Management 72:798-807.